That is to block spam sent from hijacked smtp relay servers. If Joe Schmoe installs IIS on his home box to run a website and sets up smtp with no firewall or smtp relay security, that box can be used to relay spam. Since it's a relay, the From: address will not resolve to the source IP. If you register your MX record so it matches your IP, a reverse DNS lookup will match your
user@whatever.com address to the proper IP.